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PD Dr. Johannes Schultz


 ORCID 0000-0003-4117-232X  Johannes_crop

Bonn Social Neuroscience Unit Link to the Bonn Social Neuroscience Unit


Contact Information 

Center for Economics and Neuroscience (CENs)
Zentrale wissenschaftliche Einrichtung der Universität Bonn
Am Hofgarten 8
Room: 1.008
D-53113 Bonn, Germany
Tel.: +49 (0) 228 73-8282
Fax: +49 (0) 228 73-8299
E-Mail: [Email protection active, please enable JavaScript.]
Institute for Experimental Epileptology and Cognition Research (IEECR) 
University of Bonn Medical Center | Life and Brain
Venusberg-Campus 1, Gebäude 76 
D-53127 Bonn, Germany


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Main Research Interests

  • Social perception and decision-making


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Curriculum Vitae

2019-present Acting Director (provisional), Center for Economics and Neuroscience (CENs), University of Bonn
2019-present Principal Investigator, Bonn Social Neuroscience Unit
2018-present Assistant Professor, Center for Economics and Neuroscience & Institute for Experimental Epileptology and Cognition Research, University of Bonn
2015-2018 Research Scientist and Deputy Head, Division of Medical Psychology, University Clinic Bonn
2012-2015 Lecturer (Assistant Professor), Department of Psychology, Durham University, UK
2004-2012 Research Scientist then Project Leader at Max-Planck Institute for Biological Cybernetics, Tübingen
2000-2004 PhD at the Wellcome Trust Centre for Neuroimaging, University College London, UK
1999-2000 Programme in experimental psychology and neuropsychology, Department of Psychology, University of Geneva, Switzerland
1993-1999 Medical school, Universities of Fribourg and Geneva, Switzerland


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In press



  • C, Wasserthal, S, Delis A, Schultz J, Hurlemann R, Ettinger U. (2022). Ketamine increases fronto-posterior functional connectivity during meta-perceptual confidence ratings. Behavioural Brain Research, BBR_113925
  • Lieberz J., Shamay-Tsoory S., Saporta N., Kanterman A., Gorni J., Esser T., Kuskova E., Schultz J., Hurlemann R., Scheele D. (2022). Behavioral and neural dissociation of social anxiety and loneliness. Journal of Neuroscience. DOI:


  • Coenjaerts M., Pape F., Santoso V., Grau F., Stoffel-Wagner B., Philipsen A., Schultz J., Hurlemann R., Scheele D. (2021). Sex differences in economic decision-making: Exogenous estradiol has opposing effects on fairness framing in women and men. European Neuropsychopharmacology, 50, 46-54. DOI: 10.1016/j.euroneuro.2021.04.006
  • Muthesius A., Grothey F., Cunningham C., Hölzer S., Vogeley K., Schultz J. (2021). Preserved metacognition despite impaired perception of intentionality cues in schizophrenia. Schizophrenia Research: Cognition, 100215. DOI: 10.1016/j.scog.2021.100215
  • Lehmann M., Neumann C., Wasserthal S., Schultz J., Delis A., Trautner P., Hurlemann R., Ettinger U. (2021). Effects of ketamine on brain function during metacognition of episodic memory. Neuroscience of Consciousness, 7(1): niaa028. DOI: 10.1093/nc/niaa028


  • Rramani Q., Gerhardt H.*, Grote X.*, Zhao W., Schultz J., Weber B. (2020). Do Disadvantageous Social Contexts Influence Food Choice? Evidence From Three Laboratory Experiments. Frontiers in Psychology, 11:575170. DOI: 10.3389/fpsyg.2020.575170
  • Kreuder A.-K., Scheele D., Schultz J., Hennig J., Marsh N., Dellert T., Ettinger U., Philipsen A., Babasiz M., Herscheid A., Remmersmann L., Stirnberg R., Stöcker T., Hurlemann R. (2020). Common and dissociable effects of oxytocin and lorazepam on the neurocircuitry of fear. Proceedings of the National Academy of Sciences USA, 117, 11781-11787. DOI: 10.1073/pnas.1920147117
  • Muthesius A., Hölzer S., Grothey A. F., Rubruck J., Vogeley K., Schultz J. (2020). New gambling task reveals an atypical relation between social anxiety and social avoidance in schizophrenia. Schizophrenia Bulletin Online. DOI: 10.1093/schizbullopen/sgaa013
  • Maier A. M., Gieling C., Heinen-Ludwig L., Stefan V., Schultz J., Güntürkün O., Stoffel-Wagner B., Becker B., Hurlemann R., Scheele D. (2020). Association of childhood maltreatment with interpersonal distance and social touch preferences in adulthood. American Journal of Psychiatry, 177, 37-46. DOI: 10.1176/appi.ajp.2019.19020212


  • Schultz J, Willems T., Gädeke M., Chakkour G., Franke A., Weber B., Hurlemann R. (2019). A human subcortical network underlying social avoidance revealed by risky economic choices. eLife. DOI: 10.7554/eLife.45249
  • Schultz J., Bülthoff H.H. (2019). Perceiving animacy purely from visual motion cues involves intraparietal sulcus. NeuroImage, 197, 120–132. DOI: 10.1016/j.neuroimage.2019.04.058
  • Schmidt S.J., Hurlemann R., Schultz J., Wasserthal S., Kloss C., Maier W., Meyer-Lindenberg A., Hellmich M., Muthesius-Digón A., Pantel T., Wiesner P.-S., Klosterkötter J., Ruhrmann S. (2019). Multimodal prevention of first psychotic episode through N‐acetyl‐L‐cysteine and integrated preventive psychological intervention in individuals clinically at high risk for psychosis: Protocol of a randomized, placebo‐controlled, parallel‐group trial. Early Intervention in Psychiatry, 2019, 1-12. DOI: 10.1111/eip.12781



  • Schultz J., Becker B., Preckel K., Seifert M., Mielacher C., Conrad R., Kleiman, A., Maier, W., Kendrick, K.M., Hurlemann, R. (2018). Improving therapy outcome prediction in major depression using multimodal functional neuroimaging: A pilot study. Personalized Medicine in Psychiatry, 11-12, 7-15, DOI: 10.1016/j.pmip.2018.09.001
  • Dobs K., Bülthoff I., Schultz J. (2018). Use and Usefulness of Dynamic Face Stimuli for Face Perception Studies - a Review of Behavioral Findings and Methodology. Frontiers in Psychology, 9, 403–7. DOI: 10.3389/fpsyg.2018.01355
  • Dobs K., Schultz J., Bülthoff I., Gardner J.L. (2018). Task-dependent enhancement of facial expression and identity representations in human cortex. NeuroImage, 172, 689–702. DOI: 10.1016/j.neuroimage.2018.02.013 


  • Becker B., Steffens M., Zhao Z., Kendrick K.M., Neumann C., Weber B., Schultz, J., Mehta, M.A., Ettinger, U., Hurlemann, R. (2017) General and emotion-specific neural effects of ketamine during emotional memory formation. NeuroImage, 150, 308–17. DOI: 10.1016/j.neuroimage.2017.02.049
  • Spengler F.B., Schultz J., Scheele D., Essel M., Maier W., Heinrichs M., Hurlemann, R. (2017). Kinetics and Dose Dependency of Intranasal Oxytocin Effects on Amygdala Reactivity. Biological Psychiatry, 15, 1–10. DOI: 10.1016/j.biopsych.2017.04.015
  • Hurlemann R, Scheele D, Maier W, Schultz J. (2017). Oxytocin drives prosocial biases in favor of attractive people. Behavioral and Brain Sciences, 40, e30. DOI: 10.1017/S0140525X16000807



  • Dobs K., Bülthoff I., Schultz J. (2016). Identity information content depends on the type of facial movement. Scientific Reports, 6, 34301. DOI: 10.1038/srep34301
  • David N., Skoruppa S., Gulberti A., Schultz J., Engel A.K. (2016). The Sense of Agency Is More Sensitive to Manipulations of Outcome than Movement-Related Feedback Irrespective of Sensory Modality. PLoS ONE. 2016, 11, e0161156. DOI: 10.1038/srep34301
  • de la Rosa S., Schillinger F.L., Bülthoff H.H., Schultz J., Uludag K. (2016). fMRI Adaptation between Action Observation and Action Execution Reveals Cortical Areas with Mirror Neuron Properties in Human BA 44/45. Frontiers in Human Neuroscience, 10, 78. DOI: 10.3389/fnhum.2016.00078
  • Esins J., Schultz J., Stemper C., Kennerknecht I., Bülthoff I. (2016). Face Perception and Test Reliabilities in Congenital Prosopagnosia in Seven Tests. i-Perception, 2041669515625797. DOI: 10.1177/2041669515625797



  • Kim J., Schultz J., Rohe T., Wallraven C., Lee S.-W., Bülthoff H.H. (2015). Abstract representations of associated emotions in the human brain. Journal of Neuroscience, 35, 5655–63. DOI: 10.1523/JNEUROSCI.4059-14.2015


  • Esins J., Schultz J., Wallraven C., Bülthoff I. (2014). Do congenital prosopagnosia and the other-race effect affect the same face recognition mechanisms? Frontiers in Human Neuroscience, 8, 759. DOI: 10.3389/fnhum.2014.00759
  • Esins J., Schultz, J., Bülthoff I., Kennerknecht I. (2014). Galactose uncovers face recognition and mental images in congenital prosopagnosia: The first case report. Nutritional Neuroscience, 17, 239–240. DOI: 10.1179/1476830513Y.0000000091
  • Dobs K., Bülthoff I., Breidt M., Vuong Q.C., Curio C., Schultz J. (2014). Quantifying Human Sensitivity to Spatio-Temporal Information in Dynamic Faces. Vision Research, 100, 78–87. DOI: 10.1016/j.visres.2014.04.009
  • David N.*Schultz J.*, Milne E., Schunke O., Schöttle D., Münchau A., Siegel M., Vogeley K., Engel A.K. (2014). Right Temporoparietal Gray Matter Volume Predicts Accuracy of Social Perception in Autism Spectrum Disorders. Journal of Autism and Developmental Disorders, 44:1433–1446. DOI: 10.1007/s10803-013-2008-3



  • Schultz J., Bülthoff H.H. (2013). Parametric animacy percept evoked by a single moving dot mimicking natural stimuli. Journal of Vision, 13, 15. DOI: 10.1167/13.4.15
  • Schultz J., Brockhaus M., Bülthoff H.H., Pilz K.S. (2013). What the human brain likes about moving faces. Cerebral Cortex, 23, 1167-1178. DOI: 10.1093/cercor/bhs106


2012 and earlier

  • Helbig H.B., Ernst M.O., Ricciardi E., Pietrini P., Thielscher A., Mayer K.M., Schultz J., Noppeney U. (2012). The neural mechanisms of reliability weighted integration of shape information from vision and touch. NeuroImage, 60, 1063–1072. DOI: 10.1016/j.neuroimage.2011.09.072
  • Schultz J., Lennert T. (2009). BOLD signal in intraparietal sulcus covaries with magnitude of implicitly driven attention shifts. NeuroImage, 45(4), 1314–1328. DOI: 10.1016/j.neuroimage.2009.01.012
  • Schultz J., Pilz K.S. (2009). Natural facial motion enhances cortical responses to faces. Experimental Brain Research, 194(3), 465–475. DOI: 10.1007/s00221-009-1721-9
  • Schultz J., Chuang L., Vuong Q.C. (2008). A dynamic object-processing network: metric shape discrimination of dynamic objects by activation of occipitotemporal, parietal, and frontal cortices. Cerebral Cortex, 18(6), 1302–1313. DOI: 10.1093/cercor/bhm162
  • Schultz J., Friston K.J., O'Doherty J., Wolpert D.M., Frith C.D. (2005). Activation in posterior superior temporal sulcus parallels parameter inducing the percept of animacy. Neuron, 45(4), 625–635. DOI: 10.1016/j.neuron.2004.12.052
  • Schultz J., Imamizu H., Kawato M., Frith C.D. (2004). Activation of the human superior temporal gyrus during observation of goal attribution by intentional objects. Journal of Cognitive Neuroscience, 16 (10), 1695–1705. DOI: 10.1162/0898929042947874
  • O’Doherty J., Dayan P., Schultz J., Deichmann R., Friston K.J., Dolan R.J. (2004). Dissociable roles of ventral and dorsal striatum in instrumental conditioning. Science, 304, 452–454. DOI: 10.1126/science.1094285

  * These authors contributed equally to the respective work.

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Other Articles

  • Schultz J., Hurlemann R. (2018). Characterizing variations in perceptual decision-making. Behavioral and Brain Sciences, 41: e241. DOI: 10.1017/S0140525X18001371
  • Wasserthal S., Kloss C., Hurlemann R., Schultz J. (2018). N-Acetylcystein (NAC) - eine neue Option zur frühen Bekämpfung von Schizophrenie. Der Neurologe & Psychiater, 19, 28–38. DOI: 10.1007/s15202-018-1942-5
  • Schultz J., Wasserthal S., Kloss C., Schmidt S., Ruhrmann S., Hurlemann R. (2018). Prävention einer ersten psychotischen Episode. Nervenheilkunde, 37, 313-317. DOI: 10.1055/s-0038-1651940
  • Hurlemann R., Marsh N., Schultz J., Scheele D. (2017). Oxytocin shapes the priorities and neural representations of attitudes and values. Behavioral and Brain Sciences, 40, e241. DOI: 10.1055/s-0038-1651940
  • Schultz J. (2010). Brain Imaging: Decoding Your Memories. Current Biology, 20, R269–271. DOI: 10.1016/j.cub.2010.02.001
  • Schultz J., Sebanz N., Frith C.D. (2004). Conscious will in the absence of ghosts, hypnotists, and other people. Behavioral and Brain Sciences, 27, 674–675. DOI: 10.1017/s0140525x04390151


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